Walter Hartwig, PhD

Professor, College of Osteopathic MedicineDirector of Enrollment Management and Student Success, College of Osteopathic Medicine


In my role as Director of Enrollment Management and Student Success I am responsible for stewarding our osteopathic medical students from admission through graduation. I work with each student individually to develop their path toward residency and professional clinical service. For enrollment management my goal is to communicate the TUCOM mission to prospective applicants to broaden and deepen our identity in communities most in need of access to medical education and healthcare. In the general realm of medical education I am pleased to have authored Med School Rx: Getting In, Getting Through, and Getting On With Doctoring, now in its second edition. In my faculty role I teach anatomy and embryology, and an occasional elective course in human growth and development. I am thankful that my curriculum was published as the textbook Fundamental Anatomy (Lippincott, Williams and Wilkins), in 2007. My academic background is in physical anthropology in general and primate evolution in particular. I am fortunate to have conducted fieldwork in Tanzania, Colombia, Venezuela and, most recently, Brasil. The body of my modest scholarly work has been published in two dozen peer-reviewed articles across 13 different journals, and in a dozen invited book chapters. I am privileged to have edited and produced a large reference work in primate evolution, The Primate Fossil Record (Cambridge University Press). I joined the Touro University California faculty in 1997. Prior to that I held teaching appointments at University of California at Berkeley, George Washington University, and University of Illinois-Chicago.


  • 1993 PHD Anthropology University of California-Berkeley
  • 1986 BA Anthropology University of Missouri


My published research pertains mostly to the evolution of monkeys, apes and humans, particularly in terms of the trade-offs between neural and somatic growth trajectories (1991 - 2013).  In addition to primate paleontology and life history I have enjoyed studying the history of physical anthropology (1995, 1997, 1999, 2002, 2007), and the codification of anatomy and embryology in atlas and textbook form (1990 - 1995, 2010).  

Most recently, with TUC colleagues Glenn Davis and Richard Riemer, we have been funded to study Patient Experience of Osteopathic Physician Distinction and Empathy.

Recent Publications

Books and Monographs:

2011 - Hartwig, WC  Med School Rx: Getting In, Getting Through and Getting On With Doctoring. 2nd Edition.  New York:  Kaplan Publishing.

2009 - Hartwig, WC  Med School Rx: Getting In, Getting Through and Getting On With Doctoring.  New York:  Kaplan Publishing.

2007 - Hartwig, WC  Fundamental Anatomy.  Lippincott, Williams & Wilkins.

2002 - Hartwig, WC (editor)  The Primate Fossil Record.  Cambridge:  Cambridge University Press.

Most recent publications:

2017 - Davis GE, Hartwig WC, McTighe AJ  Clinical preceptors' perceptions of empathy: the Empathy in Osteopathic Training and Education (EMOTE) study. J Am Osteopath Assoc. 117(8):503-509. doi:10.7556/jaoa.2017.100.

2015 - Rosenberger AL, Cooke SB, Halenar L, Tejedor MF, Hartwig WC, Novo NM and Munoz-Saba Y  Fossil Alouattines and the origins of Alouatta: craniodental diversity and interrelationships. In  Howler Monkeys: Examining the Evolution, Physiology, Behavior, Ecology and Conservation of the Most Widely Distributed Neotropical Primate. Ed. by M Kowalewski, P Garber, L Cortés-Ortiz, B Urbani, D Youlatos. New York: Springer Press, pp.21-54.

2013 - Hartwig, WC  Review of Ameranthropoides loysi Montandon 1929: The History of a Primatological Fraud.  Urbani, B and Viloria, AL.  Buenos Aires: Libros en Red, 2008.  The Relict Hominoid Inquiry 2:22-23.

2013 - Hartwig, WC and Barnett, A Fossil History, Zoogeography and Taxonomy of the Pitheciids: Introduction.  In Evolutionary Biology and Conservation of Titis, Sakis and Uacaris (Veiga, Barnett, Ferrari, Norconk, eds.)  Cambridge:  Cambridge University Press, pp.1-4.

2013 - Rosenberger, AL and Hartwig, WC  Primates (Lemurs, Lorises, Tarsiers, Monkeys and Apes).  In Encyclopedia of Life Sciences, 3rd edition. Chichester:  John Wiley and Sons, Ltd.  DOI: 10.1002/9780470015902.a0001572.pub3

2013 - Rosenberger, AL and Hartwig, WC  New World monkeys.  In Encylopedia of Life Sciences, 3rd edition.  Chichester:  John Wiley and Sons, Ltd. DOI: 10.1002/9780470015902.a0001562.pub3

2012 - Yavari M, Klapstein KD, Hartwig WC & Hermel E  Presence of the Functional Caspase-12 Allele in Indian Subpopulations.  Int J Immunogenetics 39:389-393.  

2011 - Hartwig WC, Rosenberger AL, Norconk MA, Young Owl M  Relative brain size, gut size and evolution in New World monkeys.  Anatomical Record 294:2207-2221.    

2011 - Hartwig, WC  Primate Evolution.  In Primates in Perspective, 2nd Edition (Campbell, Panger, Fuentes, MacKinnon and Stumpf, eds.).  Oxford University Press, pp.19-31.

2011 - Frost, SR, Rosenberger, AL, Hartwig, WC  Old World Monkeys.  In Encyclopedia of Life Sciences 2011, John Wiley and Sons, Ltd.  [DOI: 10.1002/9780470015902.a0001561.pub2]

2010 - Hartwig, WC  Review of The Human Body: An Introduction for the Biomedical and Health Sciences, by Gillian Pocock and Christopher D. Richards. Quarterly Review of Biology 85:240-241.

2008 - Hartwig, WC  Evolution on the road less traveled.  Review of Primate Evolution, by Susan Cachel.  American Journal of Primatology 70:1003-1005. 

2008 - Rosenberger AL, Halenar L, Cooke S, Hartwig WC  Morphology and evolution of the spider monkey, genus Ateles.  In Spider Monkeys:  Behavior, Ecology and Evolution of the Genus Ateles  (Campbell CJ, ed.).  Cambridge:  Cambridge University Press, pp. 19-49.

2007 - Hartwig, WC  Obituary: F. Clark Howell.  Evolutionary Anthropology16:81-82.

2007 - Hartwig, WC  Selection and the outer self.  Review of Skin:  A Natural History by Nina G. Jablonski.  Journal of Mammalian Evolution 14:200-201.

2006 - Hartwig, WC  Primate Evolution.  In Primates in Perspective (Campbell, Panger, Fuentes, MacKinnon and Bearder, eds.).  Oxford University Press, pp.11-22.

2005 - Hartwig, WC  Implications of molecular and morphological data for understanding ateline phylogeny.  International Journal of Primatology 26:999-1015.