Faculty & Staff at TUC

Walter Hartwig

Walter Hartwig

College: COM

Department: Academic Affairs

Title: Director of Enrollment Management and Student Success

Phone: (707) 638-5410


E-Mail: whartwig@touro.edu

Office: Administration & Faculty 1, Rm. 234

In my role as Director of Enrollment Management and Student Success I am responsible for stewarding our osteopathic medical students from admission through graduation.  I work with each student individually to develop their path toward residency and professional clinical service.  For enrollment management my goal is to communicate the TUCOM mission to prospective applicants to broaden and deepen our identity in communities most in need of access to medical education and healthcare.  In the general realm of medical education I am pleased to have authored Med School Rx: Getting In, Getting Through, and Getting On With Doctoring, now in its second edition.

In my faculty role I teach anatomy and embryology, and an occasional elective course in human growth and development.  I am thankful that my curriculum was published as the textbook Fundamental Anatomy (Lippincott, Williams and Wilkins), in 2007.

My academic background is in physical anthropology in general and primate evolution in particular.  I am fortunate to have conducted fieldwork in Tanzania, Colombia, Venezuela and, most recently, Brasil.  The body of my modest scholarly work has been published in two dozen peer-reviewed articles across 13 different journals, and in a dozen invited book chapters.  I am privileged to have edited and produced a large reference work in primate evolution, The Primate Fossil Record (Cambridge University Press).

I joined the Touro University California faculty in 1997.  Prior to that I held teaching appointments at University of California at Berkeley, George Washington University, and University of Illinois-Chicago.

Institution Degree Field of Study Obtained
University of California – Berkeley Ph.D Anthropology 1993
University of Missouri B.A. Anthropology 1986

Fundamentals of Osteopathic Medicine


My published research pertains mostly to the evolution of monkeys, apes and humans, particularly in terms of the trade-offs between neural and somatic growth trajectories (1991 - 2013).  In addition to primate paleontology and life history I have enjoyed studying the history of physical anthropology (1995, 1997, 1999, 2002, 2007), and the codification of anatomy and embryology in atlas and textbook form (1990 - 1995, 2010).

Research Grants

1988 Paleontological Survey of Middle Miocene Deposits at La Venta, Colombia.  Graduate Division, University of California, Berkeley

1990  Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Graduate Division, University of California, Berkeley

1991 Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Explorers Club

1991 Paleontological Survey of Tertiary Deposits along the Cordillera Merida and Orinoco Basin, Venezuela. Department of Anthropology, University of California, Berkeley

1992 Biogeography of New World Monkeys and the Pleistocene Caves of Lagoa Santa, Brazil. Sigma XI Society of Scientific Research

1992 An Interactive Image Archive System for the Study of Prehistory. National Geographic Society

1993 Workshop on Imaging Research in Physical Anthropology. LSB Leakey Foundation

1994 New Large-bodied Primates in the Pleistocene of Brazil. LSB Leakey Foundation

1998 Biogeography and Biodiversity of Pleistocene Mammals in Southeastern
Amazonia. National Geographic Society

2004 Primate Origins and Paleocommunities (POPCOM): a Global Field Test of the
Early Origins Hypothesis. Touro University – California.

2016 Patient Perspectives of Osteopathic Distinction and Physician Empathy.  (Glenn Davis and Richard Riemer, co-PIs).  American Osteopathic Association.

2011 Hartwig, WC Med School Rx: Getting In, Getting Through and Getting On With Doctoring.  2nd Edition. New York: Kaplan Publishing.

2007 Hartwig, WC Fundamental Anatomy. Baltimore: Lippincott, Williams & Wilkins

2002 Hartwig, WC (editor) The Primate Fossil Record. Cambridge: Cambridge University Press.

Partial List of Recent Articles and Chapters

2017   Davis GE, Hartwig WC, McTighe AJ Clinical preceptors' perceptions of empathy: the Empathy in Osteopathic Training and Education (EMOTE) study. J Am Osteopath Assoc. 117(8):503-509. doi:10.7556/jaoa.2017.100.

2015   Rosenberger AL, Cooke SB, Halenar L, Tejedor MF, Hartwig WC, Novo NM and Munoz-Saba Y  Fossil Alouattines and the origins of Alouatta: craniodental diversity and interrelationships. In  Howler Monkeys: Examining the Evolution, Physiology, Behavior, Ecology and Conservation of the Most Widely Distributed Neotropical Primate. Ed. by M Kowalewski, P Garber, L Cortés-Ortiz, B Urbani, D Youlatos. New York: Springer Press, pp.21-54.

2013   Hartwig, WC  Review of Ameranthropoides loysi Montandon 1929: The History of a Primatological Fraud.  Urbani, B and Viloria, AL.  Buenos Aires:  Libros en Red, 2008.  The Relict Hominoid Inquiry 2:22-23.

2013       Hartwig, WC and Barnett, A    Fossil History, Zoogeography and Taxonomy of the Pitheciids:  Introduction.  In Evolutionary Biology and Conservation of Titis, Sakis and Uacaris(Veiga, Barnett, Ferrari, Norconk, eds.)  Cambridge:  Cambridge University Press, pp.1-4.

2013   Rosenberger, AL and Hartwig, WC  Primates (Lemurs, Lorises, Tarsiers, Monkeys and Apes).  In Encylopedia of Life Sciences, 3rd edition.  Chichester:  John Wiley and Sons, Ltd.  DOI: 10.1002/9780470015902.a0001572.pub3

2013   Rosenberger, AL and Hartwig, WC  New World monkeys.  In Encylopedia of Life Sciences, 3rd edition.  Chichester:  John Wiley and Sons, Ltd. DOI: 10.1002/9780470015902.a0001562.pub3

2012    Yavari M, Klapstein KD, Hartwig WC & Hermel E  Presence of the Functional Caspase-12 Allele in Indian Subpopulations.  International Journal of Immunogenetics 39:389-393 

2011   Hartwig WC, Rosenberger AL, Norconk MA, Young Owl M  Relative brain size, gut size and evolution in New World monkeys.  Anatomical Record 294:2207-2221.             

2011   Hartwig, WC  Primate Evolution.  In Primates in Perspective, 2nd Edition (Campbell, Panger, Fuentes, MacKinnon and Stumpf, eds.).  OxfordUniversity Press, pp.19-31.

2011      Frost, SR, Rosenberger, AL, Hartwig, WC  Old World Monkeys.  In Encyclopedia of Life Sciences 2011, John Wiley and Sones, Ltd.  http://www.els.net/ [DOI: 10.1002/9780470015902.a0001561.pub2]

2010       Hartwig, WC  Review of The Human Body: An Introduction for the Biomedical and Health Sciences, by Gillian Pocock and Christopher D. Richards.  Quarterly Review of Biology 85:240-241



2008    Hartwig, WC Evolution on the road less traveled. Review of Primate Evolution, by Susan Cachel. American Journal of Primatology 70:1003-1005.

2008    Rosenberger, AL, Halenar, L, Cooke, S, and Hartwig, WC Morphology and evolution of the spider monkey, genus Ateles. In Spider Monkeys: Behavior, Encology and Evolution of the Genus Ateles (Campbell CJ, ed.). Cambridge: Cambridge University Press, pp. 19-49.

2007    Hartwig, WC Obituary: F. Clark Howell, Evolutionary Anthropology 16:81-82.

2007    Hartwig, WC Selection and the outer self. Review of Skin: A Natural History, by Nina G. Jablonski. Journal of Mammalian Evolution 14:200-201.

2006    Hartwig, WC Primate Evolution. In Primates in Perspective (Campbell, Panger, Fuentes, MacKinnon and Bearder, eds.). Oxford University Press, pp. 11-22.

2005    Hartwig, WC Implications of molecular and morphological data for understanding ateline phylogeny. International Journal of Primatology 26:999-1015.

2005    Hartwig, WC and Rosenberger, AL New World monkeys. In Encyclopedia of Life Sciences. London: Macmillan. (www.els.net/els/els/els/index.html?

2005    Hartwig, WC and Rosenberger, AL Primates. In Encyclopedia of Life Sciences. London: Macmillan. /els/els/els/index.html?sessionid=e8808eb7ecf71d4f)

2004    Hartwig, WC Review of Life’s Solution by Simon Conway Morris. American Journal of Human Biology 16:496-497

Prior Articles:

2002 Whitaker, JM, Williams, T, Rousseau, L., Rowan, RA, and Hartwig, WC A scoring system for estimating age in the foot skeleton. American Journal of Physical Anthropology 118:385-392.

2002 Hartwig, WC Introduction to the Primate Fossil Record. In The Primate Fossil Record (WC Hartwig, ed.), pp. 1-4. Cambridge: Cambridge University Press.

2002 Hartwig, WC and Meldrum, DJ Miocene platyrrhines of the northern Neotropics. In The Primate Fossil Record (WC Hartwig, ed.), pp. 175-188. Cambridge: Cambridge University Press.

2001 Hartwig, WC and Rosenberger, AL Old World monkeys. In Encyclopedia of Life Sciences. London: Macmillan. www.els.net/els/els/els/index.html? sessionid=e8808eb7ecf71d4f)

2001 Hartwig, WC Fossils, footprints and foragers: bipedalism evolving. Evolutionary Anthropology 10:3-4.

1999 Chamberlain JG and Hartwig WC Thomas Kuhn and paleoanthropology. Evolutionary Anthropology 8:42-44.

1999 Hartwig, WC Primate evolution. In The Nonhuman Primates (P Dolhinow & A Fuentes, eds). New York: Mayfield Press, pp.10-17.

1998 Masterson, TJ and Hartwig, WC Degrees of sexual dimorphism in Cebus and other New World monkeys. American Journal of Physical Anthropology 107:243-256.

1998 Hartwig, WC and Doneski, KF Evolution of the hominid hand and tool making behavior. American Journal of Physical Anthropology 106:401-404.

1998 Hartwig, WC Language origins and diversification. Evolutionary Anthropology 6:39-40.

1997 Fleagle, JG and Hartwig, WC Paleoprimatology. In History of Physical Anthropology: An Encyclopedia (F. Spencer, ed.). New York: Garland, pp.796-810.

1997 Cartelle, C and Hartwig, WC Primatologia de Sudamerica: los grandes platirrinos. Investigacion y Ciencia (April):30-32.

1997 Hartwig, WC Squirrel monkeys as animal models of pelvic relaxation. American Journal of Obstetrics and Gynecology 177:489-490.

1996 Hartwig, WC Perinatal life history traits in New World monkeys. American Journal of Primatology 40:99-130.

1996 Hartwig, WC and Cartelle, C A complete skeleton of the giant South American primate Protopithecus. Nature 381:307-311.

1996 Cartelle, C and Hartwig, WC A new extinct primate among the Pleistocene megafauna of Bahia, Brazil. Proceedings of the National Academy of Sciences 93:6405-6409.

1996 Hartwig, WC; Rosenberger, AL; Garber, PW; Norconk, MA On ateline evolution. In MA Norconck, AL Rosenberger and P Garber (eds.): Adaptive Radiations of Neotropical Primates. New York: Plenum Press, pp.427-431.

1996 Hartwig, WC Physical anthropology today and tomorrow. Evolutionary Anthropology 4:160

1996 Cartelle, C and Hartwig, WC Updating the two Pleistocene primates from Bahia, Brasil. Neotropical Primates 4:46-48.

1996 Hartwig, WC A model human biology program. Evolutionary Anthropology 5:1-2.

1996 Cartelle, C and Hartwig, WC Macacos sul-americanos: descoberta importante muda a história. Ciência Hoje 21:30-37.

1995 Hartwig, WC A giant New World monkey from the Pleistocene of Brazil. Journal of Human Evolution 28:189-196.

1995 Hartwig, WC Effect of life history on the squirrel monkey (Platyrrhini, Saimiri) cranium. American Journal of Physical Anthropology 97:435-449.

1995 Hartwig, WC Protopithecus: Rediscovering the first fossil primate. History and Philosophy of the Life Sciences 17:447-460.

1994 Hartwig, WC Patterns, puzzles and perspectives on platyrrhine origins. In Integrative Paths to the Past: Paleoanthropological Advances in Honor of F. Clark Howell (RS Corruccini and RL Ciochon, eds.). New York: Prentice-Hall, pp. 69-93.

1994 Hartwig, WC Book Review: The Cambridge Encyclopaedia of Human Evolution. Evolutionary Anthropology 2:147-149.

1994 Hartwig, WC Embryology in the context of clinical gross anatomy: a review of current texts. Clinical Anatomy 7:108-109.

1994 Hartwig, WC Setting the Future Agenda for Research on Neotropical Primates: A Symposium in Honor of Warren Kinzey. Evolutionary Anthropology 2:189-192.

1994 Hartwig, WC Book Review: Origins of Anatomically Modern Humans (MH Nitecki and DV Nitecki, eds). Quarterly Review of Biology 69:562-563.

1993 Hartwig, WC Comparative Morphology, Ontogeny and Phylogeny of the Platyrrhine Cranium. PhD Dissertation, University of California, Berkeley.

1993 Hartwig, WC and LL Sadler Visualization and the future of physical anthropology. In Milestones in Human Evolution. Perspectives in Human Evolution, Volume 6 (A. Almquist and A. Manyak, eds.). Chicago: Waveland Press, pp. 187-222.

1992 Hartwig, WC Book Review: Jablonski's Dictionary of Syndromes and Eponymic Diseases. Clinical Anatomy 5:492-493.

1991 Hartwig, WC Fractal analysis of sagittal suture morphology. Journal of Morphology 210:289-298.

1991 Hartwig, WC Book Review: Pernkopf's Anatomy, 3rd ed. (W. Platzer, ed.) Clinical Anatomy 4:63-64.

1991 Rosenberger, AL, T Setoguchi and WC Hartwig Laventiana annectens: new fossil evidence for the origin of callitrichine New World monkeys. Proceedings of the National Academy of Sciences 88:2137-2140.

1991 Rosenberger, AL, WC Hartwig, M Takai, T Setoguchi and N Shigehara Dental variability in Saimiri and the taxonomic status of Neosaimiri fieldsi, an early squirrel monkey from Colombia, South America. International Journal of Primatology 12:291-301.

1991 Hartwig, WC, T Setoguchi and AL Rosenberger The emergence of modern platyrrhine genera in the Miocene of Colombia. Proceedings of the XIIIth Congress of the International Primatological Society. Amsterdam: Elsevier, pp. 525-526.

1991 Rosenberger, AL, WC Hartwig and RG Wolff Szalatavus attricuspis, an early platyrrhine primate. Folia Primatologica 56:225-233.

1990 Hartwig, WC Dental metrics and the radiation of New World monkeys: A preliminary analysis. Kroeber Anthropological Society Papers 71/72:57-66.

1990 Hartwig, WC Book Review: The Anatomy and Biology of the Human Skeleton, by D. Gentry Steele and Claud Bramblett. Clinical Anatomy 3:151-153.


1999 Hartwig, WC Under Brazil: Mapping the Caves of Ancient Amazonia. Mercator’s World 5:24-31

2005 Hartwig, WC The intelligent design of scientific inquiry. Napa Valley Register, Guest Commentary, 11 July 2005.




1997 - present     Touro University - California

1996 - 97            University of California, Berkeley (Lecturer)

1995 - 96            George Washington University (Visiting Assistant Professor)

1993 - 95            State University of New York, Stony Brook (Postdoctoral Fellow)

1991 - 92            University of Illinois - Chicago (Instructor)

Last Updated: 4/18/22